Ovules are borne on
a)	Microsporophyll	b)	Megasporophyll	c)	Macrosporophyll	d)	Both (a) and (c)

(b)

In all cycads except the genus Cycas, the ovules are borne on megasporophylls in megastrobili, in Cycas the ovules develop on individual leaf-like megasporophylls in what is regarded as a primitive arrangement. The microspores of all cycads develop into microstrobili